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Next he appeared in a guest role on the James Cameron television series Dark Angel on Fox in 2001 as serial killer Ben/X5-493, the brother of main character Max/X5-452 (played by Jessica Alba).
His character died in the episode, but Ackles returned to the show as a series regular in the second season as Ben's sane clone, Alec/X5-494.
He also appeared on stage from June 5–10, 2007 with Lou Diamond Phillips in A Few Good Men at Casa Mañana Theatre in Fort Worth, Texas, as Lt. Ackles received strong praise for his work in this role, which was also his professional theatre debut.
In the summer of 2008, Ackles was cast in the remake of the cult film My Bloody Valentine 3D, which opened nationwide on January 16, 2009.
He also auditioned for the role of the young Clark Kent on Smallville, a role offered to actor Tom Welling instead.He also starred as the lead in the box office success My Bloody Valentine 3D and voiced Jason Todd in the popular animated film Batman: Under the Red Hood. Ackles had planned to study sports medicine at Texas Tech University and to become a physical therapist, but instead moved to Los Angeles to start his acting career.He has English, Irish, Scottish, and German ancestry.Ackles turned down the role, which was then offered to Eric Christian Olsen and the character's name was changed to "Jensen" because the producers of Tru Calling liked Ackles' name.Ackles returned to Vancouver (where Dark Angel was filmed) in 2004 to become a regular on Smallville playing the assistant football coach Jason Teague, who was also the newest romantic interest for Lana Lang (played by Kristin Kreuk).
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This role earned Jensen three Daytime Emmy nominations from 1998-2000.